Northern Atlantic Coastal Plain Pitch Pine Barrens

EVT 7355
CES203.269GNRTreeConifer
Summary
This system comprises a group of dry pitch pine woodlands and forests of deep sandy soils ranging from Cape Cod (Massachusetts) south through Long Island (New York) and the famous Pine Barrens of the New Jersey Coastal Plain, with occasional occurrences north to southernmost Maine and south to the Anacostia watershed (Maryland). The vegetation is characterized by a tree canopy of Pinus rigida with a tall-shrub layer dominated by Quercus ilicifolia and a low-shrub layer characterized by Vaccinium pallidum and/or Vaccinium angustifolium. The system is heavily influenced by fire, the composition and structure of its components varying with fire frequency. In general, tree oaks are more prevalent in those stands having a longer fire-return interval; fire frequencies of 8-10 years foster the growth of "pine plains," i.e., dwarf pine stands 1 m in height. Pine barrens with a history of more-or-less biennial burns for lowbush blueberry production may have very few trees and be characterized as sandplain grasslands. Dwarf-shrubs such as Arctostaphylos uva-ursi, Vaccinium angustifolium, Vaccinium pallidum, and Hudsonia ericoides typify the field layer of pine plains and sandplain grasslands. Schizachyrium scoparium is the most common grass (in close proximity to the coast, it may be represented by its close relative Schizachyrium littorale).

Scrub oak stands may occur without pine cover, particularly in low-lying areas that do not intersect the water table, where cold-air drainage inhibits pine growth. North of the glacial boundary, heathlands characterized by Arctostaphylos uva-ursi, Corema conradii, and Morella pensylvanica, and grasslands characterized by Schizachyrium littorale, Schizachyrium scoparium, and Danthonia spicata occur as small (or occasionally large) patches. The Pine Barrens of New Jersey are very similar in structure and composition to those north of the glacial boundary but are characterized by additional species, such as Quercus marilandica, Quercus stellata, Pyxidanthera barbulata, Leiophyllum buxifolium, and others. Where the water table is close to the surface, pitch pine lowland vegetation (described as a separate system) occurs.
Source: NatureServe Explorer
Vegetation
The uniqueness of the New Jersey Pine Barrens flora has long been recognized (Stone 1911, Harshberger 1916). More recent treatments by Forman (1979) and Buckhholz and Good (1982) have compiled much of the available information. Pinus rigida is the dominant and characteristic species of this system. It may be found in well-developed tree form or as a short-statured, shrubby ecotype. Pinus rigida may occur as the sole dominant or occur with a variety of oak species, especially Quercus marilandica, Quercus stellata, and Quercus ilicifolia. In some examples Pinus echinata may co-occur.
Source: NatureServe Explorer
Environment
This system typically occurs on deep well-drained sand deposits. In the coastal regions of the glaciated Northeast, it occurs on outwash plains and morainal deposits. In New Jersey, it occurs on Cohansey sand, which is sometimes overlain with hilltop gravel deposits.
Source: NatureServe Explorer
Dynamics
Different fire frequencies and intensities interrupt succession, accounting for variations in forest composition. Periodic severe wildfires with 40- to 100-year intervals have produced oak-pine mixtures over extensive areas of uplands, while more frequent severe fires have created mixtures of pitch pine and shrub oaks. The most frequent and severe fires have created the pine plains (Landfire 2007a).
Source: NatureServe Explorer
Threats
Fire suppression, urban and agricultural land conversion and landscape fragmentation effecting fire behavior, limiting management options, and introducing invasive plants species. On Long island, New York, pine barrens were noted to have decreased in size to 45% of their original extent (Jordan et al. 2003). The New Jersey Pine Barrens has a complex land-use history; used originally as a source of raw materials for iron smelting, ship building, transitioning over time to severe threats of conversion to residential and commercial development. Walker and Solecki (1999) documented a loss of 317 km2 of pine barrens between 1975 and 1986 to urban and agricultural uses. This represents an annual conversion rate of 0.6%, comparable to the rate of tropical deforestation conversion between 1980 and 1990.
Source: NatureServe Explorer
Distribution
This system is found in the Atlantic Coastal Plain from Delaware Bay northward through the New Jersey Coastal Plain and Long Island (New York) to Cape Cod, Massachusetts, with peripheral occurrences in Pennsylvania (historic), New Hampshire (historic), and southern Maine (Kennebunk Plains and Wells Barren).
Source: NatureServe Explorer
Ecologically Associated Plant Species

Plant species that characterize this ecosystem type, organized by vegetation stratum. These are species ecologically associated with the ecosystem, not confirmed present in any specific area.

Tree canopy

Pinus echinata, Pinus rigida, Quercus ilicifolia, Quercus marilandica

Shrub/sapling (tall & short)

Ilex glabra, Morella pensylvanica

Short shrub/sapling

Arctostaphylos uva-ursi, Corema conradii, Hudsonia ericoides, Kalmia buxifolia, Vaccinium angustifolium, Vaccinium pallidum

Herb (field)

Danthonia spicata, Pyxidanthera barbulata, Schizachyrium scoparium, Schizachyrium scoparium ssp. littorale
Source: NatureServe Ecological System assessment
Ecologically Associated Animals (5)

Animal species ecologically associated with this ecosystem type based on NatureServe assessment. These are species whose habitat requirements overlap with this ecosystem, not confirmed present in any specific roadless area.

Mammals (1)

Common NameScientific NameG-Rank
White-footed DeermousePeromyscus leucopusG5

Reptiles (2)

Common NameScientific NameG-Rank
North American RacerColuber constrictorG5
Eastern Hog-nosed SnakeHeterodon platirhinosG5

Butterflies & Moths (2)

Common NameScientific NameG-Rank
Herodias or Pine Barrens UnderwingCatocala herodias gerhardiG3T3
Common Oak MothPhoberia ingenuaG3G4
Source: NatureServe Ecological System assessment
At-Risk Species Associated with this Ecosystem (2)

Species with conservation concern that are ecologically associated with this ecosystem type. G-Rank indicates global conservation status: G1 (critically imperiled) through G5 (secure). ESA status indicates U.S. Endangered Species Act listing.

Common NameScientific NameG-RankESA Status
Herodias or Pine Barrens UnderwingCatocala herodias gerhardiG3T3--
Common Oak MothPhoberia ingenuaG3G4--
Source: NatureServe Ecological System assessment
Component Associations (14)

Plant community associations that occur within this ecological system. Associations are the finest level of the U.S. National Vegetation Classification (USNVC) and describe specific, repeating assemblages of plant species. Each association represents a distinct community type that may be found where this ecosystem occurs.

NameG-Rank
Gaylussacia baccata - Vaccinium angustifolium - Arctostaphylos uva-ursi / Schizachyrium littorale Dwarf-shrublandG3 NatureServe
Morella pensylvanica / Schizachyrium littorale - Danthonia spicata Shrub GrasslandG2 NatureServe
Pinus rigida / Carex pensylvanica WoodlandGNR NatureServe
Pinus rigida - (Pinus echinata) - Quercus stellata / Quercus (marilandica, ilicifolia) WoodlandG2 NatureServe
Pinus rigida - Quercus coccinea - Quercus falcata / (Quercus marilandica) / Gaylussacia frondosa WoodlandG2 NatureServe
Pinus rigida - Quercus coccinea / Vaccinium pallidum - (Morella pensylvanica) WoodlandGNR NatureServe
Pinus rigida - Quercus ilicifolia / Arctostaphylos uva-ursi ScrubG1 NatureServe
Pinus rigida / Quercus ilicifolia - Kalmia angustifolia / Pyxidanthera barbulata WoodlandG1 NatureServe
Pinus rigida / Quercus ilicifolia / Morella pensylvanica WoodlandG3 NatureServe
Pinus rigida - Quercus marilandica / Corema conradii ScrubG2 NatureServe
Pinus rigida / Quercus (marilandica, ilicifolia) / Pyxidanthera barbulata WoodlandG2 NatureServe
Quercus falcata - Pinus rigida - Pinus echinata / Ilex opaca WoodlandG4 NatureServe
Quercus ilicifolia - Quercus prinoides ScrubGNR NatureServe
Vaccinium angustifolium / Schizachyrium scoparium - Carex lucorum Shrub GrasslandGNR NatureServe
State Conservation Ranks (9)

Subnational conservation status ranks (S-ranks) assigned by Natural Heritage Programs in each state where this ecosystem occurs. S1 indicates critically imperiled at the state level, S2 imperiled, S3 vulnerable, S4 apparently secure, and S5 secure. An ecosystem may be globally secure but imperiled in specific states at the edge of its range.

StateS-Rank
DESNR
MASNR
MDSNR
MESNR
NHSNR
NJSNR
NYSNR
PASNR
RISNR
Methodology and Data Sources

Ecosystem classification: Ecosystems are classified using the LANDFIRE 2024 Existing Vegetation Type (EVT) layer, mapped to NatureServe Terrestrial Ecological Systems via a curated crosswalk. Each EVT is linked to the USNVC (U.S. National Vegetation Classification) hierarchy through pixel-level co-occurrence analysis of LANDFIRE EVT and NatureServe IVC Group rasters across all roadless areas.

Vegetation coverage: Coverage percentages and hectares are derived from zonal statistics of the LANDFIRE 2024 EVT raster intersected with roadless area boundaries.

Ecosystem narratives and community species: Sourced from the NatureServe Explorer API, representing professional ecological assessments of vegetation composition, environmental setting, dynamics, threats, and characteristic species assemblages.

IVC hierarchy: The International Vegetation Classification hierarchy is sourced from the USNVC v3.0 Catalog, providing the full classification from Biome through Association levels.

Component associations: Plant community associations listed as components of each NatureServe Ecological System. Association data from the NatureServe Explorer API.

State ranks: Conservation status ranks assigned by NatureServe member programs in each state where the ecosystem occurs.