Great Lakes Limestone Pavement Barrens

EVT 7409Great Lakes Alvar
CES201.721GNRHerbGrassland
Summary
Alvars are natural systems of humid and subhumid climates, centered around areas of glaciated horizontal limestone/dolomite (dolostone) bedrock pavement with a discontinuous thin soil mantle. These communities are characterized by distinctive flora and fauna with less than 60% tree cover that is maintained by associated geologic, hydrologic, and other landscape processes. In particular, all forms of alvar tend to flood each spring, then experience moderate to severe drought in summer months. They include open pavement, grassland, and shrubland/woodland types. Alvar communities occur in an ecological matrix with similar bedrock and hydrologically influenced communities. Almost all of North America's alvars occur within the Great Lakes basin, primarily in an arc along the Niagaran Escarpment from northern Lake Michigan across northern Lake Huron and eastern Ontario and northwestern New York state.
Source: NatureServe Explorer
Environment
Alvars are found near Great Lakes shores where flat limestone or dolostone bedrock pavement is exposed. Soils are shallow and discontinuous and tend to accumulate in cracks and shallow depressions in the bedrock. Where present, they are <25 cm deep. In the spring, soils are saturated or even flooded where shallow depressions occur. The thin soils dry quickly and are usually very dry by late summer.
Source: NatureServe Explorer
Dynamics
The thin soils and large changes in soil moisture during the growing season shape the vegetation of alvars. These conditions favor herbaceous species over woody species. The composition of alvars varies largely with the soil moisture from seasonal herbaceous wetlands to dry grassy areas to sparsely vegetation bedrock. Small shrublands or stunted woodlands can be found where soil accumulates (Reschke et al. 1998). Fires do not carry well on alvars in most years but they did occur with low frequency (Landfire 2007a). Woody species grow slowly on alvars, so even low frequency fires limited their abundance.
Source: NatureServe Explorer
Threats
Major threats are related to road construction, quarry development, off-road vehicle use, invasive species, and trampling of vegetation (Kost et al. 2007). Road construction results in modification of the hydrology by disrupting overland surface flows, typically flooding one side of the road and drying out the other. Road corridors and associated maintenance and off-road vehicle use facilitate the rapid introduction and expansion of invasive plants. While fire was infrequent, it was important in limiting the spread of woody species, so fire suppression activities negatively affects this system. This system recovers slowly from disturbance, so even moderate stressors can accumulate over time.
Source: NatureServe Explorer
Distribution
Alvars occur within the Great Lakes basin.
Source: NatureServe Explorer
Ecologically Associated Plant Species

Plant species that characterize this ecosystem type, organized by vegetation stratum. These are species ecologically associated with the ecosystem, not confirmed present in any specific area.

Tree canopy

Juniperus communis

Shrub/sapling (tall & short)

Dasiphora fruticosa ssp. floribunda

Herb (field)

Carex scirpoidea, Danthonia spicata, Deschampsia caespitosa, Iris lacustris, Schizachyrium scoparium, Sporobolus heterolepis
Source: NatureServe Ecological System assessment
Ecologically Associated Animals (12)

Animal species ecologically associated with this ecosystem type based on NatureServe assessment. These are species whose habitat requirements overlap with this ecosystem, not confirmed present in any specific roadless area.

Birds (4)

Common NameScientific NameG-Rank
Grasshopper SparrowAmmodramus savannarumG5
Upland SandpiperBartramia longicaudaG5
Loggerhead ShrikeLanius ludovicianusG4
Eastern TowheePipilo erythrophthalmusG5

Reptiles (1)

Common NameScientific NameG-Rank
Eastern MassasaugaSistrurus catenatusG3

Butterflies & Moths (5)

Common NameScientific NameG-Rank
Hoary ElfinCallophrys poliosG5
Mottled DuskywingErynnis martialisG3
Chryxus ArcticOeneis chryxus strigulosaG5T4
Tawny CrescentPhyciodes batesiiG3G4
Grizzled SkipperPyrgus centaureaeG5

Molluscs (1)

Common NameScientific NameG-Rank
Pleistocene CatinellaMediappendix exilisG3?

Other (1)

Common NameScientific NameG-Rank
Appalachian Grizzled SkipperPyrgus wyandotG1G2Q
Source: NatureServe Ecological System assessment
At-Risk Species Associated with this Ecosystem (6)

Species with conservation concern that are ecologically associated with this ecosystem type. G-Rank indicates global conservation status: G1 (critically imperiled) through G5 (secure). ESA status indicates U.S. Endangered Species Act listing.

Common NameScientific NameG-RankESA Status
Mottled DuskywingErynnis martialisG3--
Dwarf Lake IrisIris lacustrisG3Threatened
Pleistocene CatinellaMediappendix exilisG3?--
Tawny CrescentPhyciodes batesiiG3G4--
Appalachian Grizzled SkipperPyrgus wyandotG1G2Q--
Eastern MassasaugaSistrurus catenatusG3Threatened
Source: NatureServe Ecological System assessment
Component Associations (13)

Plant community associations that occur within this ecological system. Associations are the finest level of the U.S. National Vegetation Classification (USNVC) and describe specific, repeating assemblages of plant species. Each association represents a distinct community type that may be found where this ecosystem occurs.

NameG-Rank
Acer saccharum - Ostrya virginiana - Carya ovata - Quercus rubra Limestone WoodlandG3 NatureServe
Carya ovata / Zanthoxylum americanum / Panicum philadelphicum - Carex pensylvanica Alvar WoodlandGNR NatureServe
Danthonia spicata - Poa compressa - (Schizachyrium scoparium) GrasslandG2 NatureServe
Deschampsia cespitosa - (Sporobolus heterolepis, Schizachyrium scoparium) - Carex crawei - Packera paupercula GrasslandG2 NatureServe
Juniperus communis - (Juniperus virginiana) - Rhus aromatica - Viburnum rafinesqueanum / Oligoneuron album ShrublandG3 NatureServe
Juniperus horizontalis - Dasiphora fruticosa / Schizachyrium scoparium - Carex richardsonii Dwarf-shrublandG2 NatureServe
Juniperus virginiana / Ranunculus fascicularis WoodlandG3 NatureServe
Picea glauca - Thuja occidentalis - Juniperus communis / Iris lacustris - Carex eburnea ShrublandG1 NatureServe
Pinus banksiana - Thuja occidentalis - Picea glauca / Juniperus communis Alvar WoodlandG2 NatureServe
Sporobolus heterolepis - Schizachyrium scoparium - (Carex scirpoidea) / (Juniperus horizontalis) GrasslandG2 NatureServe
Sporobolus neglectus - Sporobolus vaginiflorus - Trichostema brachiatum - Panicum philadelphicum - (Poa compressa) Alvar GrasslandG2 NatureServe
Thuja occidentalis - Pinus banksiana / Dasiphora fruticosa / Clinopodium arkansanum Wooded GrasslandG1 NatureServe
Tortella tortuosa - Cladonia pocillum - Placynthium spp. Alvar PavementG2 NatureServe
State Conservation Ranks (4)

Subnational conservation status ranks (S-ranks) assigned by Natural Heritage Programs in each state where this ecosystem occurs. S1 indicates critically imperiled at the state level, S2 imperiled, S3 vulnerable, S4 apparently secure, and S5 secure. An ecosystem may be globally secure but imperiled in specific states at the edge of its range.

StateS-Rank
MISNR
NYSNR
OHSNR
WISNR
Methodology and Data Sources

Ecosystem classification: Ecosystems are classified using the LANDFIRE 2024 Existing Vegetation Type (EVT) layer, mapped to NatureServe Terrestrial Ecological Systems via a curated crosswalk. Each EVT is linked to the USNVC (U.S. National Vegetation Classification) hierarchy through pixel-level co-occurrence analysis of LANDFIRE EVT and NatureServe IVC Group rasters across all roadless areas.

Vegetation coverage: Coverage percentages and hectares are derived from zonal statistics of the LANDFIRE 2024 EVT raster intersected with roadless area boundaries.

Ecosystem narratives and community species: Sourced from the NatureServe Explorer API, representing professional ecological assessments of vegetation composition, environmental setting, dynamics, threats, and characteristic species assemblages.

IVC hierarchy: The International Vegetation Classification hierarchy is sourced from the USNVC v3.0 Catalog, providing the full classification from Biome through Association levels.

Component associations: Plant community associations listed as components of each NatureServe Ecological System. Association data from the NatureServe Explorer API.

State ranks: Conservation status ranks assigned by NatureServe member programs in each state where the ecosystem occurs.